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iPhone OS 3.1.3 was the last iOS software update for iPhone 2G (the original iPhone), iPod touch 1st generation and iOS 4.2.1 was the last software update for iPhone 3G, iPod touch 2nd generation users.
If you're still using them then it means that you're missing out some of the major new features introduced by Apple in iOS 4 and iOS 5 such as multitasking, folders, iCloud etc.
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